DNA-PK phosphorylation sites on Oct-1 promote cell survival following DNA damage.
Octamer transcription factor-1 (Oct-1) has recently been shown to function as a stress sensor that promotes cell survival subsequent to DNA damage. Here, we show that the survival signal imparted by Oct-1 following exposure to ionizing radiation (IR) is dependent upon DNA-dependent protein kinase (DNA-PK)-dependent phosphorylation of a cluster of ... 13 specific ser/thr residues within the N-terminal transcriptional regulatory domain of Oct-1. Although IR treatment did not affect the recruitment of Oct-1 to the histone H2B promoter, the recruitment of RNA polymerase II, TATA-binding protein and histone H4 acetylation were strongly reduced, consistent with a decrease in Oct-1 transcriptional regulatory potential following IR exposure. Ser/Thr-Ala substitution of 13 sites present in Oct-1 transcriptional regulatory domain eliminated Oct-1 phosphorylation subsequent to IR exposure. Further, these substitutions prevented Oct-1 from rescuing the survival of IR-treated Oct-1-/- murine embryonic fibroblasts, providing a direct link between DNA-PK-dependent phosphorylation and the contribution of Oct-1 to cell survival. These results implicate Oct-1 as a primary effector in a DNA-PK-dependent cell survival pathway that is activated by double-stranded DNA breaks.
Mesh Terms:
3T3 Cells, Amino Acid Sequence, Amino Acid Substitution, Animals, Binding Sites, Blotting, Western, Cell Line, Cell Line, Tumor, Cell Survival, DNA Damage, DNA-Activated Protein Kinase, Dose-Response Relationship, Radiation, Histones, Humans, Mice, Mice, Knockout, Molecular Sequence Data, Octamer Transcription Factor-1, Phosphorylation, Promoter Regions, Genetic, Protein Binding, Serine, Threonine, Transfection
3T3 Cells, Amino Acid Sequence, Amino Acid Substitution, Animals, Binding Sites, Blotting, Western, Cell Line, Cell Line, Tumor, Cell Survival, DNA Damage, DNA-Activated Protein Kinase, Dose-Response Relationship, Radiation, Histones, Humans, Mice, Mice, Knockout, Molecular Sequence Data, Octamer Transcription Factor-1, Phosphorylation, Promoter Regions, Genetic, Protein Binding, Serine, Threonine, Transfection
Oncogene
Date: Jun. 07, 2007
PubMed ID: 17213819
View in: Pubmed Google Scholar
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