Tlg2p, a yeast syntaxin homolog that resides on the Golgi and endocytic structures.
Intracellular membrane fusion events in eukaryotic cells are thought to be mediated by protein-protein interactions between soluble N-ethylmaleimide-sensitive factor attachment protein receptor (SNARE) complex proteins. We have identified and analyzed a new yeast syntaxin homolog, Tlg2p. Tlg2p is unique among known syntaxin family proteins in possessing a sizeable hydrophilic domain ... of 63 amino acids that is C-terminal to the membrane spanning region and nonessential for Tlg2p function. Tlg2p resides on the endosome and late Golgi by co-localization with an endocytic intermediate and co-fractionation with markers for both endosomes and late Golgi. Cells depleted for Tlg2p missort a portion of carboxypeptidase Y and are defective in endocytosis. In addition, we report that Tlg2p forms a SEC18-dependent SNARE complex with Snc2p, a vesicle SNARE known to function in Golgi to plasma membrane trafficking. Based on these findings we propose that Tlg2p is a t-SNARE that functions in transport from the endosome to the late Golgi and on the endocytic pathway.
Mesh Terms:
Adenosine Triphosphatases, Amino Acid Sequence, Carboxypeptidases, Cathepsin A, Cell Compartmentation, Endocytosis, Fungal Proteins, Golgi Apparatus, Macromolecular Substances, Membrane Fusion, Membrane Proteins, Membrane Transport Proteins, Molecular Sequence Data, Mutagenesis, Insertional, Qa-SNARE Proteins, R-SNARE Proteins, Receptors, Cytoplasmic and Nuclear, Saccharomyces cerevisiae, Saccharomyces cerevisiae Proteins, Sequence Alignment, Sequence Homology, Amino Acid, Vesicular Transport Proteins
Adenosine Triphosphatases, Amino Acid Sequence, Carboxypeptidases, Cathepsin A, Cell Compartmentation, Endocytosis, Fungal Proteins, Golgi Apparatus, Macromolecular Substances, Membrane Fusion, Membrane Proteins, Membrane Transport Proteins, Molecular Sequence Data, Mutagenesis, Insertional, Qa-SNARE Proteins, R-SNARE Proteins, Receptors, Cytoplasmic and Nuclear, Saccharomyces cerevisiae, Saccharomyces cerevisiae Proteins, Sequence Alignment, Sequence Homology, Amino Acid, Vesicular Transport Proteins
J. Biol. Chem.
Date: May. 08, 1998
PubMed ID: 9565594
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