Pyk2 and FAK regulate neurite outgrowth induced by growth factors and integrins.
Integration of signalling pathways initiated by receptor tyrosine kinases and integrins is essential for growth-factor-mediated biological responses. Here we show that co-stimulation of growth-factor receptors and integrins activates the focal-adhesion kinase (FAK) family to promote outgrowth of neurites in PC12 and SH-SY5Y cells. Pyk2 and FAK associate with adhesion-based complexes ... that contain epidermal growth factor (EGF) receptors, through their carboxy- and amino-terminal domains. Expression of the C-terminal domain of Pyk2 or of FAK is sufficient to block neurite outgrowth, but not activation of extracellular-signal-regulated kinase (ERK). Moreover, activation and autophosphorylation of Pyk2/FAK, as well as of effectors of their adhesion-targeting domains, such as paxillin, are important for propagation of signals that control neurite formation. Thus, Pyk2/FAK have important functions in signal integration proximal to integrin/growth-factor receptor complexes in neurons.
Mesh Terms:
Animals, Cytoskeletal Proteins, Enzyme Activation, Epidermal Growth Factor, Focal Adhesion Kinase 1, Focal Adhesion Kinase 2, Focal Adhesion Protein-Tyrosine Kinases, Growth Substances, Humans, Insulin, Insulin-Like Growth Factor I, Integrins, Mitogen-Activated Protein Kinase 3, Mitogen-Activated Protein Kinases, Nerve Growth Factor, Neurites, PC12 Cells, Paxillin, Phosphoproteins, Protein-Tyrosine Kinases, Rats, Receptor, Epidermal Growth Factor, Signal Transduction, Tumor Cells, Cultured
Animals, Cytoskeletal Proteins, Enzyme Activation, Epidermal Growth Factor, Focal Adhesion Kinase 1, Focal Adhesion Kinase 2, Focal Adhesion Protein-Tyrosine Kinases, Growth Substances, Humans, Insulin, Insulin-Like Growth Factor I, Integrins, Mitogen-Activated Protein Kinase 3, Mitogen-Activated Protein Kinases, Nerve Growth Factor, Neurites, PC12 Cells, Paxillin, Phosphoproteins, Protein-Tyrosine Kinases, Rats, Receptor, Epidermal Growth Factor, Signal Transduction, Tumor Cells, Cultured
Nat. Cell Biol.
Date: Sep. 01, 2000
PubMed ID: 10980697
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