Naf1 p is a box H/ACA snoRNP assembly factor.
Box H/ACA small nucleolar ribonucleoprotein particles (snoRNPs) contain four essential proteins, Cbf5p, Gar1p, Nhp2p, and Nop10p, each of which, with the exception of Gar1p, is required for box H/ACA snoRNA accumulation. Database searches identified a novel essential protein, which we termed Naf1p, with a region of homology to the RNA-binding ... domain of Gar1p and other features in common with hnRNP-like proteins. Naf1p is localized to the nucleus and is not a stable component of the H/ACA snoRNPs, but it is required for the accumulation of all box H/ACA snoRNAs. This requirement is not at the level of snoRNA transcription initiation or termination. Naf1 p shows in vitro RNA-binding activity and also binds directly to Cbf5p and Nhp2p. Naf1p was shown to bind to the CTD in vivo in a two-hybrid assay, and the phosphorylated CTD, but not the nonphosphorylated CTD, was shown to precipitate tagged Naf1p from a cell lysate. We propose that Naf1 p is recruited to the CTD of RNA polymerase II and binds to nascent box H/ACA snoRNAs promoting snoRNP assembly.
Mesh Terms:
Amino Acid Motifs, Amino Acid Sequence, Binding Sites, Cell Nucleus, Fungal Proteins, Hydro-Lyases, Microtubule-Associated Proteins, Molecular Sequence Data, Nuclear Proteins, RNA Polymerase II, RNA Processing, Post-Transcriptional, RNA, Ribosomal, RNA, Small Nucleolar, RNA-Binding Proteins, Recombinant Proteins, Ribonucleoproteins, Small Nuclear, Ribonucleoproteins, Small Nucleolar, Saccharomyces cerevisiae Proteins, Sequence Homology, Amino Acid, Transcription, Genetic, Yeasts
Amino Acid Motifs, Amino Acid Sequence, Binding Sites, Cell Nucleus, Fungal Proteins, Hydro-Lyases, Microtubule-Associated Proteins, Molecular Sequence Data, Nuclear Proteins, RNA Polymerase II, RNA Processing, Post-Transcriptional, RNA, Ribosomal, RNA, Small Nucleolar, RNA-Binding Proteins, Recombinant Proteins, Ribonucleoproteins, Small Nuclear, Ribonucleoproteins, Small Nucleolar, Saccharomyces cerevisiae Proteins, Sequence Homology, Amino Acid, Transcription, Genetic, Yeasts
RNA
Date: Dec. 01, 2002
PubMed ID: 12515383
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