Negative regulation of mitosis by wee1+, a gene encoding a protein kinase homolog.
Fission yeast wee1- mutants initiate mitosis at half the cell size of wild type. The wee1+ activity is required to prevent lethal premature mitosis in cells that overproduce the mitotic inducer cdc25+. This lethal phenotype was used to clone wee1+ by complementation. When wee1+ expression is increased, mitosis is delayed ... until cells grow to a larger size. Thus wee1+ functions as a dose-dependent inhibitor of mitosis, the first such element to be specifically identified and cloned. The carboxy-terminal region of the predicted 112 kd wee1+ protein contains protein kinase consensus sequences, suggesting that negative regulation of mitosis involves protein phosphorylation. Genetic evidence indicates that wee1+ and cdc25+ compete in a control system regulating the cdc2+ protein kinase, which is required for mitotic initiation.
Mesh Terms:
Amino Acid Sequence, Base Sequence, DNA Restriction Enzymes, Genes, Genes, Fungal, Genes, Lethal, Genes, Regulator, Mitosis, Protein Kinases, Saccharomycetales, Schizosaccharomyces, Sequence Homology, Nucleic Acid
Amino Acid Sequence, Base Sequence, DNA Restriction Enzymes, Genes, Genes, Fungal, Genes, Lethal, Genes, Regulator, Mitosis, Protein Kinases, Saccharomycetales, Schizosaccharomyces, Sequence Homology, Nucleic Acid
Cell
Date: May. 22, 1987
PubMed ID: 3032459
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